Home > Staff > Academic > A Prof Muthama Muasya

Associate Professor Muthama Muasya

Phone: 021 650 3725 
Room No.: 3.18.2 (HW Pearson Building)








My career began in Kenya, where I was a Research Scientist at the National Museums of Kenya (1993-2008). I have a BSc and MPhil from Moi University (Kenya), PhD from University of Reading (UK), and postdoctoral stints at Royal Botanic Gardens Kew (UK), Rutgers University (USA) and KU Leuven (Belgium). I am a Senior Lecturer (since 2006) teaching courses in Biodiversity and Evolutionary Biology and my research is predominantly on Angiosperm Systematics.

Research Interests

  • Cypereceae: taxonomy and conservation; phylogenetics and biogeography; genome evolution
  • The taxonomy, speciation and macroevolution of the Cape flora with current focus on Cyperaceae and Fabaceae
  • Wetland diversity and sustainable extraction of macrophytes
  • The origin, diversity and utilization of the African savanna biome

Ongoing research collaborations

Floral ontogeny and inflorescence development in Cyperaceae
Co-researchers: Dr A. Vrijdaghs (KU Leuven), Prof P. Geotghebeur and research team (Ghent University); webpagina's/alex_vrijdaghs.html

Cyperaceae floristics, monography and family level systematics
Co-researchers: Dr D. Simpson (Royal Botanic Gardens Kew), Prof J. Bruhl (University of New England);

Genome evolution in Cyperales
Co-researchers: Prof P. Bures (Masaryk University), and team of Cyperologists from Australia, Belgium, America.

Fabaceae evolution (including role of rhizobia and ecology) and molecular systematics
Co-researchers: Dr S. Chimphango (UCT), Prof C. Stirton (UCT), Prof JI Sprent (University of Dundee), Dr E James (James Hutton Institute)

Research students

Postdoctoral (current)
B. Lemaire: Evolutionary history of bacterial root symbiosis in Cape legumes

Postdoctoral (recent)
B Gerhke: Are the mode of chromosome rearrangement and inflorescence structure influencing the evolution of Cariceae? An analysis of Schoenoxiphium (Cyperaceae)

Doctoral (current)
P. Maistry (co-supervised with S Chimphango, A. Valentine): Effect of enhanced P acquisition strategies on symbiotic N2-fixation in indigenous legumes adapted to the low P soils of the Cape Floristic Region.

K. Bauters (co-supervised with P. Goetghebeur, Ghent University): A morphological, phylogenetic and taxonomical study of the genus Scleria.

Doctoral (recently graduated)
T. Terer (co-supervised with L. Treist Vrije Universiteit Brussel): Sustainable harvesting and resilience of Cyperus papyrus resources in wetlands communities of Kenya. 2011.

G. Mwachala (co-supervised with E. Fischer, University of Koblez): Systematic and ecological studies of Dracaena (Dracaenaceae) in tropical Africa. 2005.

Masters by dissertation (current)
D. Basic (co-supervised with S Chimphango): Phosphorus uptake in cluster root and non-cluster root forming species indigenous to the Cape Floristic Region

R. Munyai (co-supervised with S Chimphango): Are old monotypic lineages ecologically and genetically impoverished?

O. Dlodlo (co-supervised with S Chimphango): characterization of rhizobia isolated from nodules of indigenous legumes and their effect on host plant distribution in the Cape Floristic Region, South Africa,

J. Viljoen (co-supervised with T Verboom): Edaphic Ecology and the Evolution of the Sedges in the Cape Flora

A. Tshiila (co-supervised with S Chimphango): Systematics of Ficinia clade (Cyperaceae)

Masters by dissertation (recently graduated)
M. Dludlu (co-supervised with S Chimphango): Systematics of Psoraleeae. 2010

F. Corry (co-supervised with J. Day): Macrophytes in wetland health. 2011.

P. Musili (co-supervised with L. Newton, Kenyatta University): Taxonomy of Cyperus rotundus complex. 2007.

G. Ngugi (co-supervised with L. Newton, Kenyatta University): Ethnobotany of Kiang'ombe Hill. 2007.

A. Ahmed (co-supervised with E. Omino Jomo Kenyatta University of Agriculture & Technology): Taxonomy of some Commiphora in E. Africa. 2004.

P. Segawa, (co-supervised with E. Kakudidi, Makerere University): The diversity and distribution of sedges in selected Important Bird Areas in Uganda. 2002.

Publications include:


Gehrke B., Vrijdaghs A., Smets E., Muasya A.M. 2012. Unisexual flowers as a robust synapomorphy in Cariceae (Cyperaceae)? Evidence for bisexual flowers in Schoenoxiphium. South African Journal of Botany 78: 150–158.

Muasya A.M., Reynders M., Goetghebeur P., Simpson D .A. & Vrijdaghs A. 2012. Dracoscirpoides (Cyperaceae) — a new genus from Southern Africa, its taxonomy and floral ontogeny. South African Journal of Botany 78: 104–115.

Muasya A.M., J.-A. Viljoen, Stirton C.H., N.A. Helme. 2012. Two new leafless species of Ficinia (Cypereae, Cyperaceae) from the Greater Cape Floristic Region of South Africa. South African Journal of Botany 79: 96–101.

Reynders M., Vrijdaghs A., Larridon I., Huygh W., Leroux O., Muasya A.M. & Goetghebeur P. 2012. Gynoecial anatomy and development in Cyperoideae (Cyperaceae, Poales): congenital fusion of carpels facilitates evolutionary modifications in pistil structure. Plant Ecology and Evolution 145: 96–125.

Terer T, Muasya A.M., Triest L. 2012. Effects of harvesting Cyperus papyrus in undisturbed wetlands, Lake Nairvasha, Kenya. Hydrobiologia 680: 135–148.

Terer T, Muasya A.M., Dahdouh-Guebas F., Ndiritu G.G., Triest L. 2012. Integrating local ecological knowledge and management practices of an isolated semi-arid papyrus swamp (Loboi, Kenya) into a wider conservation framework. Journal of Environmental Management 93: 71–84.


Davies T.J., Smith G.F., Bellstedt D.U., Boatwright J.S., Bytebier B., Cowling R.M., Forest F.l., Harmon L.J., Muasya A.M., Schrire B.D., Steenkamp Y., van der Bank M. & Savolainen V. (2011) Extinction Risk and Diversification Are Linked in a Plant Biodiversity Hotspot. PLoS Biol, 9, e1000620 [Download pdf Document].

Larridon I., Reynders M., Huygh W., Bauters K., Van de putte K., Muasya A.M., Boeckx P., Simpson D.A., Vrijdaghs A. & Goetghebeur P. 2011. Affinities in C3 Cyperus (Cyperaceae) revealed using molecular phylogenetic data and carbon isotope analysis. Botanical Journal of the Linnean Society 167: 19–46.

Larridon I., Reynders M., Huygh W., Bauters K., Vrijdaghs A., Leroux O., Muasya A.M., Simpson D.A., & Goetghebeur P. 2011. Taxonomic changes in C3 Cyperus (Cyperaceae) supported by molecular data, morphology, embryography, ontogeny and anatomy. Plant Ecology and Evolution 144: 327–356.

Larridon I., Reynders M., Huygh W., Muasya A. M., Govaerts R., Simpson D.A. & Goetghebeur P. 2011. Nomenclature and typification of generic and subdivisional names in the Cyperus clade (Cyperaceae): 2. Subdivisional names in Cyperus. Taxon 60: 868-884.

Muasya A.M., Larridon I., Reynders M., Huygh W., Goetghebeur P., Cable S., Simpson D.A. & Gehrke B. 2011. The Cyperaceae in Madagascar show high diversification in high altitude forest and wetland habitats. Plant Ecology and Evolution 144: 357–362.

Ngugi G.W., Newton L.E., Muasya A.M. 2011. The Contribution of Forest Products to Dryland Household Economy: The case of Kiang’ombe hill forest, Kenya. Ethnobotany Research & Applications19: 163-180.

Reynders M., Huygh W., Larridon I., Muasya A.M., Govaerts R., Simpson D.A. & Goetghebeur P. 2011. Nomenclature and typification of generic and subdivisional names in the Cyperus clade (Cyperaceae): 3. Subdivisional names in satellite genera. Taxon 60: 885-895.

Sievers S., Muasya A.M. 2011, Identification of the sedge Cladium mariscus subsp. jamaicense and its possible use in the Middle Stone Age at Sibudu, KwaZulu-Natal. Southern African Humanities 23: 77–86. 

Stirton C.H., Clark V.R, Barker N.P., Muasya A.M. (2011). Psoralea margaretiflora (Psoraleeae, Fabaceae): A new species from the Sneeuberg Centre of Floristic Endemism, Eastern Cape, South Africa. Phytokeys 5: 31–38 [Download pdf Document].

Stirton C.H., Muasya A.M. 2011. Aspalathus abbottii (Fabaceae: Crotalarieae), a new species from KwaZulu-Natal, South Africa. South African Journal of Botany77: 675–679.

Vrijdaghs A., Reynders M., Muasya A.M., Larridon I., Goetghebeur P.,  Smets E.F. 2011. Morphology and development of spikelets and flowers in Cyperus and Pycreus (Cyperaceae). Plant Ecology and Evolution. 144: 44–63.

Muasya A.M., Viljoen J., Stirton C.H. 2011. Isolepis namaquana (Cypereae, Cyperaceae), a new endemic species from the winter rainfall area of South Africa. South African Journal of Botany 77: 55–58.

Warren B.H., Bakker F.T., Bellstedt D.U., Bytebier B., ClaSZen-Bockhoff R., Dreyer L.L., Edwards D., Forest F. Galley C., Hardy C.R. Linder H.P., Muasya A.M., Mummenhoff K., Oberlander K.C., Quint M., Richardson J.E., Savolainen V., Schrire B.D., van der Niet T., Verboom G.A., Yesson C., Hawkins J.A. 2011. Consistent phenological shifts in the making of a biodiversity hotspot: the Cape flora. BMC Evolutionary Biology 11:39. doi:10.1186/1471-2148-11-39.


Gehrke B., Martín-Bravo S., Muasya A.M., & Luceño M. 2010. Monophyly, phylogenetic position and the role of hybridization in Schoenoxiphium Nees (Cariceae, Cyperaceae). Mol. Phylogenet. Evol. 56: 380–392.

Huygh W., Larridon I., Reynders M., Muasya A.M., Govaerts R., Simpson D .A. & Goetghebeur P. 2010. Nomenclature and typification of names of genera and subdivisions of genera in Cypereae (Cyperaceae): 1. Names of genera in the Cyperus clade. Taxon 59: 1883–1890.

Muasya A.M. & de Lange P. 2010. Ficinia spiralis (Cyperaceae) a new genus and combination for Desmoschoenus spiralis. New Zealand Journal of Botany 48: 31-39.

Muasya A.M., Harvey H., Cheek M., Tah K. & Simpson D. A. 2010. Coleochloa domensis (Cyperaceae), a new epiphytic species from Cameroon. Kew Bulletin 65: 323325.

Muasya A.M., Musili P.M., Vrijdaghs A. 2010. Kyllinga mbitheana (Cyperaceae)—description, floral ontogeny and pollen micromorphology of a new species from Kenya. Journal of East African Natural History 99: 65–75.

Moore G., Smith G.F., Figueiredo E., Demissew S., Lewis G., Schrire B., Rico L., van Wyk B., Irtifaq Ali S., Ameka G. K., Atahuachi M., Bandyopadhyay S., BarkerN., Beentje H., Bruneau A., Bytebier B., Craven L., Crisp M., Cunningham A., Dold T., Fagg C., Fortuna A.N., Fortunato R. H., Friis I., Garcia F., Glazewski J.1, Grobler A., Hall M., Herendeen P., Hughes C., Huntley B. J., Kalema J., Kativu S., Klitgaard B., Knees S., Kwembeya E., Lakshminarasimhan P., Landrum L., Linder P., Luckow M., Mackinder B., van der Maesen J., Miller A. G., Moll E., Muasya A.M., Mwachala G., Neale S., Nee M. H., Pathak M. K., Predeep S. V., Sanjappa M., Smith P., Srivastava S. K., Stefano R. D., Stirton C., Timberlake J., Torke B. M., Varma S. K., Wieringa J., Wojciechowski M. 2010. Acacia, the 2011 Nomenclature Section in Melbourne, and beyond. Taxon 59: 1188–1195.

Sieben E.J.J., Morris C.D., Kotze D.C. & Muasya A.M. 2010. Changes in plant form and function across altitudinal and wetness gradients in the wetlands of the Maloti-Drakensberg, South Africa. Plant Ecology 207: 107-119.

Vrijdaghs A., Reynders M., Larridon I., Muasya A.M., Smets E. & Goetghebeur P. 2010. Spikelet structure and development in Cyperoideae (Cyperaceae): a monopodial general model based on ontogenetic evidence. Annals of Botany105: 555–571.

Wilkin P., Burrows J., Burrows S., Muasya A.M. & van Wyk E. 2010. A critically endangered new species of yam (Dioscorea strydomiana Wilkin, Dioscoreaceae) from Mpumalanga, South Africa. Kew Bulletin 65: 421433.


Besnard, G., Muasya A.M., Russier, F., Roalson, E.H., Salamin, N. & Christin, P.A.  2009. Phylogenomics of C4 Photosynthesis in Sedges (Cyperaceae): Multiple Appearances and Genetic Convergence. Molecular Biology and Evolution 26: 1909–1919.

Hoenselaar K., Muasya A.M. & Simpson D. A. 2009. Fuirena bidgoodae, a new species of Cyperaceae from East Africa. Kew Bulletin 64: 685–687.

Muasya A.M., Simpson D.A., Verboom G.A., Goetghebeur P., Naczi R.F.C., Chase M.W. & Smets E. 2009. Phylogeny of Cyperaceae based on DNA sequence data: current progress and future prospects. Botanical Review 75: 2-21 [ Download pdf Document ].

Muasya A.M., Vrijdaghs A., Simpson D.A., Chase M.W., Goetghebeur P. & Smets E. 2009. What is a genus in Cypereae: phylogeny, character homology assessment and generic circumscription. Botanical Review 75: 52-66 [ Download pdf Document ].

Nagels, A., Muasya A.M., Huysman, S., Vrijdaghs A., Smets E. & Vinckier S. 2009. Palynological diversity in Cyperaceae - a search for informative pollen and orbicule characters. Plant Systematics & Evolution 277: 117-142.

Verboom G.A., Archibald J.K., Bakker F.T., Bellstedt D.U., Conrad F., DreyerL.L., Forest F., Galley C., Goldblatt P., Henning J.F., Mummenhoff K., Linder H.P., Muasya A.M., Oberlander K.C., Savolainen V., Snijman D.A., van der Niet T., Nowell T.L. 2009. Origin and diversification of the Greater Cape flora: Ancient species repository, hot-bed of recent radiation, or both? Molecular Phylogenetics & Evolution 51: 44-53.

Vrijdaghs A., Muasya A.M., Goetghebeur P., Caris P., Nagels A. & Smets E. 2009. A floral ontogenetic approach to homology questions within the Cyperoideae (Cyperaceae). Botanical Review 75: 30-51 [Download pdf Document].

Wilkin P., Muasya A.M., Banks H., Furness C.A., Vollesen K., Weber O., & Demissew S. 2009. A new species of yam (Dioscorea) from Kenya: pollen morphology, conservation status, and speciation. Systematic Botany 34: 652-659.


Christin P.A. Salamin N., Muasya A.M., Roalson E.H., Russier F. & G. Besnard. 2008. Genetic convergence exposes determinants of Rubisco higher efficiency in C4 plants. Molecular Biology and Evolution 25: 2361-2368 .

Knox E.B., Muasya A.M. & N. . 2008 The predominantly South American clade of Campanulaceae subfam. LobelioideaeSystematic Botany 33:462-468.

Knox E.B., Heenan, P.B., Muasya A.M. & B.G. Murray 2008. Phylogenetic position and relationships of Lobelia glaberrima (Lobeliaceae), a new alpine species from southern South Island ( New Zealand). New Zealand Journal of Botany 46: 77-85.


Muasya A.M. , Simpson D.A. & Smets E. 2007. Isolepis levynsiana, a new name for Cyperus tenellus (Cyperaceae). Novon 17: 59.

Mwachala G., Fischer E. & Muasya A.M. 2007. A new species of Dracaena L. (Dracaenaceae-Ruscaceae) from Mt Kupe and the Bakossi Mts, Cameroon. Kew Bulletin 62: 613-616.

Simpson, D.A. & Muasya, A.M. 2007. Phylogeny of Cyperaceae: an overview. In: Barbos, L.M. & Santos Jr, N.A. (eds). A botânica no Brasil: pesquisa, ensini e políticas públicas ambientais, pp. 308-310. Sociedade Botânica do Brasil, Sao Paulo.

Simpson D.A., Muasya A.M., Alves M., Bruhl J. J., Dhooge S., Chase M. W., Furness C. A., Ghamkhar K., Goetghebeur P., Hodkinson T. R., Marchant A. D., Nieuborg R., Reznicek A. A., Roalson E. H., Smets E., Starr J. R., Thomas W. W., Wilson K. L. & Zhang X. 2007. Phylogeny of Cyperaceae based on DNA sequence data - a new rbcL analysis. Aliso 23: 72-83.


Knox, E. B., Muasya A. M., and P. B. Phillipson. 2006. The Lobeliaceae originated in southern Africa. In S. A. Ghazanfar & H. J. Beentje (eds.), Taxonomy and Ecology of African Plants, their Conservation and Sustainable Use. Proceedings of the XVIIth AETFAT Congress, Addis Ababa, Ethiopia. Pp. 215–227. Royal Botanic Gardens, Kew, U.K.

Masinde, P. S. & A. M. Muasya. 2006. Centenary of the East African Herbarium: 1902 –2002. In S. A. Ghazanfar & H. J. Beentje (eds.), Taxonomy and Ecology of African Plants, their Conservation and Sustainable Use. Proceedings of the XVIIth AETFAT Congress, Addis Ababa, Ethiopia. Pp. 699–707. Royal Botanic Gardens, Kew, U.K.

Muasya A.M. , Simpson D.A. & Smets E. 2006. Isolepis tenella, a new combination in Cyperaceae. Novon 16: 89–90.

Vrijdaghs A., Goetghebeur P., Smets E. & Muasya A.M. (2006). The floral scales in Hellmuthia (Cyperaceae, Cyperoideae) and Paramapania (Cyperaceae, Mapanioideae): an ontogenetic study . Annals of Botany 98: 619 – 630.


Muasya A.M. (2005). Ficiniaanysbergensis and F.esterhuyseniae (Cyperaceae), two new species from the Cape Floristic Region of South Africa. South African Journal of Botany 71: 197–200.

Muasya A.M. & Simpson D.A. 2005. Validation of Isolepis sect. Proliferae (Cyperaceae). Kew Bulletin 60: 159.

Parnell J.A.N., Simpson D.A., Chayamarit K., Boonthavikoon T., Boyce P.C., Chantaranothai P., De Wilde B., Jebb M.H.P., Muasya A.M., Paton A.J., Pendry C.A., Pooma R., Sudee S. & Wilkin, P. 2005. The Bangkok Forest Herbarium, Royal Botanic Gardens Kew and Trinity College Dublin plant collecting trips in Thailand 1995–2002. Thai Forest Bulletin (Botany) 33: 145–156.

Simpson D.A., Muasya A.M., Chayamarit K., Parnell J.A.N., Suddee, S., Wilde B.D.E, Jones M.B., Bruhl, J.J. & Pooma, R. 2005. Khaosokia caricoides, a new genus and species of Cyperaceae from Thailand. Botanical Journal of the Linnean Society 149: 357–364.

Vrijdaghs A., Goetghebeur P., Muasya A.M., Caris P. & Smets E. (2005). Floral ontogeny in Ficinia Schrad. and Isolepis R.Br. (Cyperaceae), with focus on the nature and origin of the gynophore. Annals of Botany 96: 1247 – 1262.


Ashley G.M., Maitima Mworia, J., Muasya A.M., Owen R.B., Driese S.G., Hover V.C., Renaut R.W., Goman M.F., Mathai, S., & Blatt S.H. 2004. Sedimentation and recent history of a freshwater wetland in a semi-arid environment, Loboi Swamp, Kenya, East Africa. Sedimentology 51: 1–23.

Muasya A.M. & Simpson D.A. 2004. Cyperus kituiensis (Cyperaceae), a new species from Kenya. Kew Bulletin 59: 247–249.

Muasya A.M. , Hover V.C., Ashley G.M., Owen R.B., Goman M.F. & Kimeli M. 2004. Diversity and distribution of macrophytes in a freshwater wetland, Loboi Swamp (Rift Valley) Kenya. Journal of East African Natural History 91: 39–47.

Owen R.B., Renaut R.W., Hover V.C., Ashley G.M. & Muasya A.M. 2004. Swamps, springs and diatoms: wetlands of the semi-arid Bogoria-Baringo Rift, Kenya. Hydrobiologia 518: 59–78.

Simpson D.A. & Muasya A.M. 2004. Three new species of Cyperus (Cyperacea) from eastern and southern Africa. Kew Bulletin 59: 593–598.

Ssegawa P., Kakudidi E., Muasya A.M. & Kalema J. 2004. Diversity and distribution of sedges on multivariate environmental gradients. African Journal of Ecology 42(1): 21–33.

Vrijdaghs A., Goetghebeur P, Muasya A.M., Smets E. & Caris P. 2004. The nature of the perianth in Fuirena (Cyperaceae). South African Journal of Botany. 70: 587 –594.


Dhooge, S., Goetghebeur P. & Muasya A.M. 2003. Zameioscirpus , a new genus of Cyperaceae from South America. Plant Systematics & Evolution 243(1-2): 73-84.

Parnell J.A.N., Simpson D.A., Moat J., Kirkup D.W., Chantaranothai P., Boyce P.C., Bygrave P., Dransfield S., Jebb M.H.P., Macklin J., Meade C., Middleton D.J., Muasya A.M., Prajaksood A., Pendry C.A., Pooma R., Sudee S. & Wilkin, P. 2003. Plant collecting spread and densities; their potential on biogeographic studies in Thailand. Journal of Biogeography 30: 193–209.

Simpson D.A., Furness C.A., Hodkinson T.R., Muasya A.M. & Chase M.W. 2003. Phylogenetic relationships in Cyperaceae subfamily Mapanioideae inferred from pollen and plastid DNA sequence data. American Journal of Botany 1071–1086.


Ashley G.M., Goman M., Hover V.C., Owen R.B., Renaut R.W. & Muasya A.M. 2002. Artesian blister wetlands, a perennial water resource in the semi-arid Rift Valley of East Africa. Wetlands 22: 686–695.

Malombe I., Kelbessa E., & Muasya A.M. 2002. A taxonomic study of the Blepharisedulis complex (Acanthaceae) in Eastern Africa. Journal of East African Natural History 91: 81–90.

Muasya A.M. 2002. Kyllingiella simpsonii (Cyperaceae), a new species from Tropical Africa. Kew Bulletin 57: 997–999.

Muasya A.M. & Simpson D.A. 2002. A monograph of the genus Isolepis R.Br. (Cyperaceae). Kew Bulletin 57: 257–362.

Muasya A.M. , Simpson D.A. & Chase M.W. 2002. Phylogenetic relationships in Cyperus s.l. (Cyperaceae) inferred from plastid DNA sequence data. Botanical Journal of the Linnean Society 138: 145–153.


Muasya A.M. , Simpson D.A. & Chase M.W. 2001. Generic relationships and character evolution in Cyperus s.l. (Cyperaceae). Systematics & Geography of Plants 71: 539–544.

Muasya A.M. , Simpson D.A., Chase M.W. & Culham A. 2001. A phylogeny of Isolepis (Cyperaceae) inferred using plastid rbcL and trnL–F sequence data. Systematic Botany 26: 342–353.

Simpson D.A., Muasya A.M. & Cafferty S. 2001. Typification of Scirpus fluitans and Scirpus setacea (Cyperaceae). Kew Bulletin 56: 1011–1012.


Muasya A.M. , Bruhl J.J., Simpson D.A., Chase M.W. & Culham A. 2000. Suprageneric phylogeny of Cyperaceae: a combined analysis. In K.L. Wilson & D.A. Morrison (eds.), Monocots: Systematics and Evolution, 593–601. CSIRO, Melbourne. ISBN 0 643 06437 0.

Muasya A.M. , Simpson D.A., Chase M.W. & Culham A. 2000. Phylogenetic relationships within the heterogeneous Scirpus sensu lato. (Cyperaceae) inferred from rbcL and trnL–F sequence data. In K.L. Wilson & D.A. Morrison (eds.), Monocots: Systematics and Evolution, 610–614. CSIRO, Melbourne. ISBN 0 643 06437 0.

Muasya A.M. , Simpson D.A. & Goetghebeur P. 2000. New combinations in Trichophorum, Scirpoides and Ficinia (Cyperaceae). Novon 10: 132–133.


Muasya A.M. 1998. A synopsis of Fuirena (Cyperaceae) for the Flora Tropical East Africa. Kew Bulletin 53: 187–202.

Muasya A.M. , Simpson D.A., Chase M.W. & Culham A. 1998. An assessment of suprageneric phylogeny in Cyperaceae using rbcL DNA sequences. Plant Systematics & Evolution 211: 257–271.

Book article

Muasya A.M. 2002. Isolepis R.Br. In B. Vald?s, M. Rejdali, A. Achhal El Kadmiri, S.L. Jury & J.M. Montserrat (eds.), Checklist of Vascular Plants of N Morocco with Identification Keys, 746. CSIC, Madrid. ISBN 84 00 08073 4